WP-Members 3.5.5.1 is a bug fix release for version 3.5.5.
This version patches a bug in the function that checks if a user has access to a post. When the membership required for access has both a role requirement and an expiration, the function returns true if the user has the role, regardless of their expiration date.
This is not a common combination, so in most cases, an update may not be necessary. However, if you have a defined membership that uses both expiration and a role requirement, this patch is a necessary update.