WP-Members 3.5.5.1 is a bug fix release for version 3.5.5.
This version patches a bug that does not correctly check access if a post has a membership requirement that has both an expiration date and a required role.
This is not a common combination, so in most cases, an update may not be necessary. However, if you have a defined membership that uses both expiration and a role requirement, this patch is a necessary update.